Q: Which team reached the FA Cup Final at Wembley even though
they lost a game (that was not abandoned or replayed) in an
earlier round in the same season?
A: Charlton Athletic in 1945/46. They lost 2-1 to Fulham in the
third round, but for that season only each round up to and
including the quarter-finals was played over two legs. Charlton
won the first leg of that third round tie 3-1, so went through on
aggregate.
Sometimes this question is phrased as which team won the FA Cup having lost a game earlier in the competition. However, that is wrong as Derby County beat Charlton in the final, and did not lose a game in any of the earlier rounds.
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